Course information provided by the 2026-2027 Catalog.
Considers explanations for pattern of diversity and the apparent good fit of organisms to the environment. Topics include the diversity of life, the genetics and developmental basis of evolutionary change, processes at the population level, evolution by natural selection, modes of speciation, long-term trends in evolution, origin of humans.
Enrollment Information Priority given to: first-years, sophomores, and transfer students.
Distribution Requirements (BIO-AG, BSC-AG, OPHLS-AG), (BIO-AS)
Exploratory Studies (CU-SBY)
Last 4 Terms Offered 2026FA, 2026SP, 2025FA, 2025SP
Learning Outcomes
Understand the underlying causal principles of evolutionary diversification.
Apply these principles to understand historical and contemporary evolutionary scenarios.
Identify ~100 core taxa in the tree of life, their characteristics, and understand the relationships among them.
Use basic conceptual and analytical tools to describe complex relationships within the tree of life.
Become familiar with a number of experimental and synthetic approaches to analyzing and discovering evolutionary processes (microevolution) and establishing evolutionary patterns (macroevolution).
Write and discuss knowledgeably about the dimensions of evolutionary issues that require decisions in our society.
Regular Academic Session.Choose one lecture and one discussion.
Combined with:
BIOEE 1780
Credits and Grading Basis
4 Credits
Stdnt Opt(Letter or S/U grades)
Class Number & Section Details
5297BIOEE 1780 LEC 001
Meeting Pattern
T
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5300BIOEE 1780 DIS 201
Meeting Pattern
T
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5301BIOEE 1780 DIS 202
Meeting Pattern
T
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5302BIOEE 1780 DIS 203
Meeting Pattern
T
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5303BIOEE 1780 DIS 204
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5304BIOEE 1780 DIS 205
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5305BIOEE 1780 DIS 206
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5306BIOEE 1780 DIS 207
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5307BIOEE 1780 DIS 208
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5308BIOEE 1780 DIS 209
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5309BIOEE 1780 DIS 210
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5310BIOEE 1780 DIS 211
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5311BIOEE 1780 DIS 212
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5312BIOEE 1780 DIS 213
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5313BIOEE 1780 DIS 214
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Syllabi:
Regular Academic Session.Choose one lecture and one discussion.
Combined with:
BIOEE 1780
Credits and Grading Basis
4 Credits
Stdnt Opt(Letter or S/U grades)
Class Number & Section Details
5298BIOEE 1780 LEC 002
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5314BIOEE 1780 DIS 215
Meeting Pattern
T
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5315BIOEE 1780 DIS 216
Meeting Pattern
T
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5316BIOEE 1780 DIS 217
Meeting Pattern
T
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5317BIOEE 1780 DIS 218
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5318BIOEE 1780 DIS 219
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5319BIOEE 1780 DIS 220
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5320BIOEE 1780 DIS 221
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5321BIOEE 1780 DIS 222
Meeting Pattern
W
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5322BIOEE 1780 DIS 223
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5323BIOEE 1780 DIS 224
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5324BIOEE 1780 DIS 225
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5325BIOEE 1780 DIS 226
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5326BIOEE 1780 DIS 227
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Class Number & Section Details
5327BIOEE 1780 DIS 228
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Syllabi:
Regular Academic Session.Choose one lecture and one discussion.
Combined with:
BIOEE 1780
Credits and Grading Basis
5 Credits
Stdnt Opt(Letter or S/U grades)
Class Number & Section Details
5375BIOEE 1780 LEC 003
Meeting Pattern
T
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Students interested in the 5-credit option must complete the application first day of class.
Class Number & Section Details
5328BIOEE 1780 DIS 229
Meeting Pattern
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Instructor Consent Required (Add)
Syllabi:
Regular Academic Session.Choose one lecture and one discussion.
Combined with:
BIOEE 1780
Credits and Grading Basis
5 Credits
Stdnt Opt(Letter or S/U grades)
Class Number & Section Details
5299BIOEE 1780 LEC 004
Meeting Pattern
R
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
Additional Information
Instruction Mode: In Person
Students interested in the 5-credit option must complete the application first day of class.
Instructor Consent Required (Add)
Class Number & Section Details
5329BIOEE 1780 DIS 230
Meeting Pattern
Jan 25 - May 11, 2027
Instructors
Drake, A
Greischar, M
To be determined. There are currently no textbooks/materials listed, or no textbooks/materials
required, for this section. Additional information may be found on the syllabus provided by your professor.
For the most current information about textbooks, including the timing and options for purchase, see the
Cornell Store.
